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Paleoparasitological Studies on Mummies of the Joseon Dynasty, Korea

Min Seo1, Adauto Araujo2, Karl Reinhard3, Jong Yil Chai4, Dong Hoon Shin5
The Korean Journal of Parasitology 2014;52(3):235-242.
Published online: June 26, 2014

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Corresponding author (bbbenji@naver.com; cuteminjae@gmail.com)
• Received: September 16, 2013   • Revised: May 12, 2014   • Accepted: May 14, 2014

© 2014, Korean Society for Parasitology and Tropical Medicine

This is an Open Access article distributed under the terms of the Creative Commons Attribution Non-Commercial License (http://creativecommons.org/licenses/by-nc/3.0/) which permits unrestricted non-commercial use, distribution, and reproduction in any medium, provided the original work is properly cited.

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Paleoparasitological Studies on Mummies of the Joseon Dynasty, Korea
Korean J Parasitol. 2014;52(3):235-242.   Published online June 26, 2014
DOI: https://doi.org/10.3347/kjp.2014.52.3.235
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Paleoparasitological Studies on Mummies of the Joseon Dynasty, Korea
Korean J Parasitol. 2014;52(3):235-242.   Published online June 26, 2014
DOI: https://doi.org/10.3347/kjp.2014.52.3.235
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Paleoparasitological Studies on Mummies of the Joseon Dynasty, Korea
Image Image Image Image Image Image
Fig. 1 Parasitological sampling from Korean mummy. (A) Making a hole for endoscopic exploration for Korean mummy. Coprolites could be collected during endoscopic examination. (B) Very well preserved feces could be obtained during the dissection of Gangneeung mummy.
Fig. 2 (A) False-positivity due to the contamination of sample could be ruled out by examination of the archaeological site's surface soils. The negative control sample could be obtained from the surface soil (black dot) adjacent to the tomb. (B) When the parasitological samples were collected from geological strata, negative control could be obtained from the nearby surface soil (black dot). (C) Joseon mummies discovered tightly wrapped in layers of clothing and other textiles. Researchers wearing sterilized gowns, masks, head caps, and gloves tried to remove the textile before paleoparasitological sampling. (D) The paleoparasitological study on a child mummy of Joseon period, found in Yangju-gun.
Fig. 3 Microscopic examination of the sample. Trichuris eggs (A) could be observed in coprolite sample of Yangju mummy. Bar=100 µm. Eggs of Trichuris trichiura (B) and Ascaris lumbricoides (C) in coprolite samples from Korean mummy. Bar=10 µm (B & C). (A) is from Yangju mummy, (B) from Mungeyong mummy, and (C) from SN1-2 case. Clonorchis sinensis egg (D) discovered in the geological strata sample. Since there were no parasite eggs in negative controls, the results could be regarded as authentic. Bar=10 µm. Scanning electron microscopic view of Trichuris trichiura (E) and Ascaris lumbricoides eggs (F) in the coprolite from a child mummy, Yangju. Bars=15 µm.
Fig. 4 (A) A larva of Strongyloides stercoralis recovered from the Gongju mummy. (B) A larva of Trichostrongylus sp. recovered from the Gongju mummy. (C) An egg of Metagonimus yokogawai recovered from the Hadong-1 mummy. Bar=10 µm. (D) An egg of Gymnophalloides seoi recovered from the Hadong-1 mummy. Bar=10 µm.
Fig. 5 Ectopic paragonimiasis in a Joseon female case. In the mummified liver sample, many Paragonimus westermani eggs were found. Bar=50 µm.
Fig. 6 (A) The places where 15 Korean mummies were discovered. Paleoparasitological studies have been done on the coprolites from each Korean mummy. 1, Yongin; 2, Jinju; 3, Sapgyo; 4, Hadong-2; 5, Hadong-1; 6, Sacheon; 7, Gangneung; 8, Dangjin; 9, Mungyeong; 10, Waegwan, 11, PJ-SM; 12, Seocheon; 13, Yangju; 14, SN (Sinnae); 15, GJ (Gongju). (B) Gymnophalloides seoi infection was identified in 2 cases of Korean mummies from (1) Sapgyo and (2) Hadong (indicated by blue dots). In a 2001 national survey, G. seoi infection was confined to small islands or counties in the southwestern corner of the Korean peninsula (areas shaded in red).

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Paleoparasitological Studies on Mummies of the Joseon Dynasty, Korea
No. Area Estimated date Date of excavation Sex Parasite eggs from ancient mummies
Al Tt Hk Ev Cs Pw My Gs Ss Tr 1 Yongin 15-16C 2007-8 Female + + + 2 Jinju 15-16C 2010-7 Female + + + 3 Sapgyo 16C 2011-2 Male + + + + 4 Hadong-2 16-17C 2009-6 Female + 5 Hadong-1 17C 2006-4 Female + + + 6 Sacheon 1620-1630 2011-10 Female + 7 Gangneung 1622 2007-10 Male + 8 Dangjin 1633 2008-8 Male + + + 9 Mungyeong 1647 2010-4 Female + + 10 Waegwan 1648 2008-10 Male + + 11 PJ SM 1699 2008-5 Female + + 12 Seocheon 17C 2007-2 Female + + 13 Yangju 1640-1700 2001-11 Male + + + 14 SN2-19-1 1765 ±10 2007-8 Male + + 15 GJ1-2 17-18C 2008-4 Male + + + + + 16 SN 1-2a 1605-1733 2007-8 Undecidable + 17 SN3-7-1a 16-17C 2007-9 Undecidable + 18 SN2-19-2a 1755±10 2007-8 Undecidable + + Total 18 samples of mummies (15) & ancient remains (3); 9 (Al), 14 (Tt) 0 (Hk), 1 (Ev), 5 (Cs), 5 (Pw), 3 (My), 2 (Gs), 1 (Ss), 1 (Tr) Species Target region Consensus sequence (5ʹ→3ʹ) Ref. Ascaris lumbricoides 18sRNA ATGGCCTATAAAGGTGAAACCGCGAACGGCTCATTACAACAGCTATTATATACTTGATCTTGATATCCTACGTGGATAACTGTGGTAATTCTAGAGCTAATACATGCACCAAAGCTCCGATTTTCTGACGAGCGCATCTATTAGATTAAAACCAATCGGGTTTCGGCCCGTCAATT [23] cyt b TTTTGACTTATGGTTGGAATTTTGGTAGTATGTTGGGTATGGTTTTAGGTTTTCAGATTTTGACTGGTACTTTTTTGGCTTTTTATTATTCTAATGAT Trichuris trichiura SSUrRNA GGAACGATCGAGCCGTACATAAGTTCGGCTAAATCTATTGGAGGGCAAGTCTGGTGCCAGCAGCCGCGGTAATTCCAGCTCCAATAGCGTATATTAAAGTTGCTGCGGTTAAACCGCTCGTAGTTGGATTGCGGATGTCGACGACGGTCGTCCTAAGCAGGAGTCGTTCCGTCGCTCGTCACCTGTTCGATCAAGATTGCCCTGGATGCTCTTCAGTGAGTGTCCTTGGCGACTTGAAAGTTTACTTTGAGAAAATGAGAGCGCTCAAGGCAAGCCGTAGT [24] Paragonimus westermani ITS2 ATATTGCGGCCACGGGTTAGCCTGTGGCCACGCCTGTCCGAGGGTCGGCTTATAAACTATCGCGACGCCCAAAAAGTCGCGGCTTGGGTTTTGCCAGCTGGCGTGATCTCCCCAATCTGGTCTTGTGCCTGTGGGGTGCCAGATCTGTGGCGTTTCCCTAACATACTCGGGCGCACCCACGTTGCGGCTGAAAGCCTTGACGGGGATGTGGCAACGGAATCGTGGCTCAGTGAATGATTTATGTGCGCGTTCCGCTGTCCTGTCTTCATCTGTGGTTTATGTTGCGCGTGGTCTGCTTTCGATGCTGACCTACGTATGTGCCATGTGGTTCATTCTCCTGACCTCGGATCAGACGTGAGTACC [40] COI Fail Clonorchis sinensis ITS1 GTCTTGCAGCATTGTCTGCCTAGGGCGGAGCGATCCTAGTTCCGTCATGTTCTACATGTATGTTCCGCATGCATGCTGCAGCGTTGTCTGCCTACGGTGGAGCGTCTCTAGTTCTACCAATTCCTGGCTATACCTGGCACGTGTACCCAATATATATGATGTGCCTACGTACAGTCGCGTTTCGGCAGGGTGCCTACCCGTCTGATGCTCTCGGTATGCTCGCTTCCGTTGGTGGCCAGTCCATATTGGGGGTGACGGGATGTGCTGTCAGAATGGACAGTGCTAGGCTTAATGAGTGGGCATGATGTGTCTCGAGCTACGGCTCACCCACCGCCCTGATGTTGTTGTTCATTTCAAACCGTTTTACACTGTTAAAGTGTTTCAGGTTGGCCTGGCCTGGCCTGACTGGCTGGCCGGCTTGTCTCACTGCCCCGACATGCACCCGGTGTTCTACACTGGACTGCATGTGCAGTCGCCCGGCGGTGCCTTATC [39] ITS2 TGTCGATGAAGAGCGCAGCCAACTGTGTGAATTAATGCGAACTGCATACTGCTTTGAACATCGACATCTTGAACGCATATTGCGGCCATGGGTTTGCCTGTGGCCACGCCTGTCCGAGGGTCGGCTTATAAACTATCACGACGCCCAAAAAGTCGTGGCTTGGGTCTTGCCAGCTGGCATGATTTCCCCACACAATTGTGTGTATGTGTGTGGGGTGCCGGATCTATGGCTTTTCCCCAATGTGCCGGACGCAACCATGTCTGGGCTGACTGCCTAGATGAGGGGGTGGCGGCGGAGTCGTGGCTCAATTGTTGTTATTGTTGTGAATGTGCGCGCTCCGTTGTTGGTCCTTTGTCTTTGGTTGAGGCTTCAGTATTGGCAATGCATTCGATGCAAATCTGTTTTGCACCGGTCGGTGCTTAACTTTCCTGACCTCGGATCAGACGTGATT COI TATGATTAGTCACATTTGTACTACTTTAACAGGTAAAGATTCGTTGTTTGGTTATGGGGGCTTGGTGTTGGCTATGTTTGCTATAGTTTGTCTGGGTAGGGTGGTTTGAGCTCATCATATGTTTACTGTTGGGCTGGATTTGGGGACTGCTGTTTTTTTTAGCT
Table 1. Parasite eggs discovered in mummies or ancient remains of Korea according to geographical areas

Al, Ascaris lumbricoides; Tt, Trichuris trichiura; Hk, hookworm; Ev, Enterobius vermicularis; Cs, Clonorchis sinensis; Pw, Paragonimus westermani; My, Metagonimus yokogawai; Gs, Gymnophalloides seoi; Ss, Strongyloides stercoralis; Tr, Trichostrongylus sp.

Ancient remains only.

Table 2. DNA sequences of ancient parasite eggs from No. 12 (A. lumbricoides), No. 18 (T. trichiura), No. 4 (P. westermani), and No. 10 (C. sinensis) mummies

18SrRNA, 18S small subunit ribosomal RNA; cyt b, cytochrome b; SSUrRNA, small subunit ribosomal RNA; ITS2, internal transcribed spacer 2; COI, cytochrome c oxidase subunit I; ITS1, internal transcribed spacer 1.

Table 1.

Table 2.

PHD : Parasites, Hosts and Diseases

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