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Original Article

Seasonal Distribution of Ticks in Four Habitats near the Demilitarized Zone, Gyeonggi-do (Province), Republic of Korea

Sung Tae Chong1, Heung Chul Kim1, In-Yong Lee2, Thomas M. Kollars3, Alfredo R. Sancho4, William J. Sames5, Joon-Seok Chae6, Terry A. Klein7,
The Korean Journal of Parasitology 2013;51(3):319-325.
Published online: June 30, 2013

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Corresponding author (terry.a.klein2.civ@mail.mil)
• Received: March 20, 2013   • Revised: April 21, 2013   • Accepted: April 30, 2013

© 2013, Korean Society for Parasitology and Tropical Medicine

This is an Open Access article distributed under the terms of the Creative Commons Attribution Non-Commercial License (http://creativecommons.org/licenses/by-nc/3.0/) which permits unrestricted non-commercial use, distribution, and reproduction in any medium, provided the original work is properly cited.

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Seasonal Distribution of Ticks in Four Habitats near the Demilitarized Zone, Gyeonggi-do (Province), Republic of Korea
Korean J Parasitol. 2013;51(3):319-325.   Published online June 30, 2013
DOI: https://doi.org/10.3347/kjp.2013.51.3.319
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Seasonal Distribution of Ticks in Four Habitats near the Demilitarized Zone, Gyeonggi-do (Province), Republic of Korea
Korean J Parasitol. 2013;51(3):319-325.   Published online June 30, 2013
DOI: https://doi.org/10.3347/kjp.2013.51.3.319
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Seasonal Distribution of Ticks in Four Habitats near the Demilitarized Zone, Gyeonggi-do (Province), Republic of Korea
Image Image
Fig. 1 Geographical location of tick collection sites at US installations and US and ROK operated training sites conducted near the demilitarized zone, Gyeonggi Province, April-October 2004-2005.
Fig. 2 Seasonal distribution of 3 tick species collected from 4 different habitats at US and ROK operated military training sites and installations and private lands located near the DMZ, Republic of Korea, from April through October 2004 and 2005 (GR, grasses and portions of herbaceous and crawling vegetation; DF, deciduous forest; CF, conifer forest; MF, mixed forest).

Fig. 1

Fig. 2

Seasonal Distribution of Ticks in Four Habitats near the Demilitarized Zone, Gyeonggi-do (Province), Republic of Korea
Habitats (No. cates*) Species Months
Mean (Total) Numbers Percent by habitats (%) Apr May Jun Jul Aug Sep Oct Grass (n = 98) Haemaphysalis longicornis Larvae 4.6 (37) 4.4 (88) 1.6 (26) 25.3 (557) 137.3 (1,648) 428.2 (5,138) 24.6 (197) 78.5 (7,691) 58.6 Nymphs 184.4 (1,475) 29.2 (592) 123.1 (1,970) 29.3 (645) 7.6 (84) 1.8 (22) 0.0 (0) 48.9 (4,788) 36.5 Adults 0.4 (3) 1.5 (30) 10.6 (170) 15.4 (339) 7.8 (94) 0.3 (3) 0.0 (0) 6.5 (639) 4.9 Subtotal 189.4 (1,515) 35.5 (710) 135.4 (2,166) 70.0 (1,541) 152.2 (1,826) 430.3 (5,163) 24.6 (197) 133.9 (13,118) 98.9 Haemaphysalis flava Larvae 0.0 (0) 0.0 (0) 0.0 (0) 0.0 (0) 0.0 (0) 0.4 (5) 0.4 (3) 0.1 (9) 16.1 Nymphs 0.0 (0) 1.1 (22) 0.2 (3) 0.2 (5) 0.0 (0) 0.0 (0) 1.1 (9) 0.4 (39) 69.6 Adults 0.0 (0) 0.2 (4) 0.1 (1) 0.0 (0) 0.0 (0) 0.1 (1) 0.3 (2) 0.1 (8) 14.3 Subtotal 0.0 (0) 1.3 (26) 0.3 (4) 0.3 (6) 0.0 (0) 0.5 (6) 1.8 (14) 0.6 (56) 0.4 Ixodes nipponensis Larvae 0.0 (0) 0.0 (0) 0.0 (0) 0.6 (13) 2.6 (31) 0.0 (0) 0.0 (0) 0.4 (44) 46.8 Nymphs 0.6 (5) 0.8 (15) 0.2 (3) 0.0 (1) 0.1 (1) 0.1 (1) 0.1 (1) 0.3 (27) 28.7 Adults 0.4 (3) 0.5 (9) 0.6 (10) 0.0 (0) 0.0 (0) 0.0 (0) 0.1 (1) 0.2 (23) 24.5 Subtotal 1.0 (8) 1.2 (24) 0.8 (13) 0.6 (14) 2.7 (32) 0.1 (1) 0.3 (2) 1.0 (94) 0.7 Total grasses Larvae 4.6 (37) 4.4 (88) 1.6 (26) 26.0 (571) 139.9 (1,679) 428.6 (5,143) 25.0 (200) 79.0 (7,744) 58.4 Nymphs 185.0 (1,480) 31.5 (629) 123.5 (1,976) 29.6 (651) 7.1 (85) 1.9 (23) 1.3 (10) 49.5 (4,854) 36.6 Adults 0.8 (6) 2.2 (43) 11.3 (181) 15.4 (339) 7.8 (94) 0.3 (4) 0.4 (3) 2.5 (670) 5.0 Total 190.4 (1,523) 38.0 (760) 136.4 (2,183) 71.0 (1,561) 154.8 (1,858) 430.8 (5,170) 26.6 (213) 135.4 (13,268) 100.0 Deciduous forests (n=68) Haemaphysalis longicornis Larvae 0.0 (0) 8.3 (33) 0.3 (4) 2.9 (23) 27.6 (442) 56.5 (226) 2.9 (58) 11.6 (786) 57.5 Nymphs 19.3 (77) 18.0 (72) 17.1 (205) 2.0 (16) 1.1 (17) 0.8 (3) 0.0 (0) 5.7 (390) 28.6 Adults 0.3 (1) 2.0 (8) 6.3 (75) 10.5 (75) 1.4 (22) 0.0 (0) 0.0 (0) 2.8 (190) 13.9 Subtotal 19.5 (78) 28.3 (113) 23.7 (284) 15.4 (123) 30.1 (481) 57.3 (229) 2.9 (58) 20.1 (1,366) 34.3 Haemaphysalis flava Larvae 0.0 (0) 0.0 (0) 0.0 (0) 0.0 (0) 116.5 (1,864) 3.3 (13) 12.9 (258) 31.4 (2,138) 89.2 Nymphs 4.3 (17) 3.3 (13) 4.8 (57) 0.6 (5) 0.6 (9) 1.0 (4) 6.4 (127) 3.4 (232) 9.7 Adults 0.0 (0) 0.8 (3) 0.2 (2) 0.0 (0) 0.4 (6) 0.0 (0) 0.9 (17) 0.4 (28) 1.1 Subtotal 4.3 (17) 4.0 (16) 4.9 (59) 0.6 (5) 117.4 (1,879) 4.3 (17) 20.1 (402) 35.2 (2,395) 60.2 Ixodes nipponensis Larvae 0.0 (0) 0.0 (0) 0.0 (0) 3.3 (26) 9.2 (147) 1.5 (6) 0.3 (5) 2.7 (184) 83.6 Nymphs 0.5 (2) 1.5 (6) 1.8 (21) 0.1 (1) 0.1 (2) 0.0 (0) 0.0 (0) 0.5 (32) 14.6 Adults 0.0 (0) 0.5 (2) 0.1 (1) 0.0 (0) 0.0 (0) 0.0 (0) 0.1 (1) 0.1 (4) 1.8 Subtotal 0.5 (2) 2.0 (8) 1.8 (22) 3.4 (27) 9.3 (149) 1.5 (6) 0.3 (6) 3.2 (220) 5.5 Total deciduous forests Larvae 0.0 (0) 8.3 (33) 0.3 (4) 6.1 (49) 153.3 (2,453) 61.3 (245) 16.1 (321) 45.8 (3,105) 78.0 Nymphs 24.0 (96) 22.8 (91) 23.6 (283) 2.8 (22) 1.8 (28) 1.8 (7) 6.4 (127) 9.6 (654) 16.4 Adults 0.3 (1) 3.3 (13) 6.5 (78) 10.5 (84) 1.8 (28) 0.0 (0) 0.9 (18) 3.3 (222) 5.6 Total 24.3 (97) 34.3 (137) 30.4 (365) 19.4 (155) 156.8 (2,509) 63.0 (252) 23.3 (466) 58.5 (3,981) 100.0 Conifer forests (n= 27) Haemaphysalis longicornis Larvae 0.0 (0) 0.0 (0) 0.0 (0) 0.2 (2) 5.9 (47) 0.8 (3) 0.4 (3) 1.0 (55) 42.0 Nymphs 0.5 (1) 1.5 (34) 1.1 (13) 0.5 (6) 0.6 (5) 0.0 (0) 0.0 (0) 1.1 (59) 45.0 Adults 0.0 (0) 0.0 (0) 0.4 (5) 0.9 (11) 0.1 (1) 0.0 (0) 0.0 (0) 0.3 (17) 13.0 Subtotal 0.5 (1) 1.5 (34) 1.5 (18) 1.6 (19) 6.6 (53) 0.8 (3) 0.4 (3) 2.4 (131) 9.2 Haemaphysalis flava Larvae 0.0 (0) 0.0 (0) 0.0 (0) 4.3 (52) 38.8 (310) 3.3 (13) 0.9 (7) 7.1 (382) 46.9 Nymphs 4.5 (9) 11.5 (167) 2.3 (28) 2.7 (32) 1.9 (15) 0.5 (2) 16.3 (130) 7.1 (383) 47.0 Adults 1.0 (2) 1.8 (16) 0.6 (7) 0.7 (8) 1.4 (11) 0.5 (2) 0.5 (4) 0.9 (50) 6.1 Ixodes nipponensis Subtotal 5.5 (11) 13.3 (183) 2.9 (35) 7.7 (92) 42.0 (336) 4.3 (17) 17.6 (141) 15.1 (815) 57.4 Larvae 0.0 (0) 0.0 (0) 0.0 (0) 35.8 (430) 2.4 (19) 0.3 (1) 0.3 (2) 8.4 (452) 95.4 Nymphs 0.0 (0) 0.3 (6) 0.3 (4) 0.2 (2) 0.8 (6) 0.0 (0) 0.0 (0) 0.3 (18) 3.8 Adults 0.0 (0) 0.1 (1) 0.0 (0) 0.2 (2) 0.0 (0) 0.0 (0) 0.1 (1) 0.1 (4) 0.8 Subtotal 0.0 (0) 0.3 (7) 0.3 (4) 36.2 (434) 3.1 (25) 0.3 (1) 0.4 (3) 8.8 (474) 33.4 Total conifer forests Larvae 0.0 (0) 0.0 (0) 0.0 (0) 40.3 (484) 47.0 (376) 4.3 (17) 1.5 (12) 16.5 (889) 62.6 Nymphs 5.0 (10) 13.3 (207) 3.8 (45) 3.3 (40) 3.3 (26) 0.5 (2) 16.3 (130) 8.5 (460) 32.4 Adults 1.0 (3) 1.8 (17) 1.0 (12) 1.8 (21) 1.5 (12) 0.5 (2) 0.6 (5) 1.3 (72) 5.0 Total 6.0 (12) 15.2 (224) 4.8 (57) 45.4 (545) 51.8 (414) 5.3 (21) 18.4 (147) 26.3 (1,420) 100.0 Mixed forests (n = 52) Haemaphysalis longicornis Larvae 0.0 (0) 0.0 (0) 0.0 (0) 0.1 (1) 85.0 (340) 0.0 (0) 0.0 (0) 6.6 (341) 84.2 Nymphs 1.1 (18) 1.3 (10) 0.8 (3) 0.1 (2) 3.0 (12) 0.0 (0) 0.0 (0) 0.9 (45) 11.1 Adults 0.0 (0) 0.0 (0) 0.3 (1) 0.5 (9) 2.3 (9) 0.0 (0) 0.0 (0) 0.4 (19) 4.7 Subtotal 1.1 (18) 1.3 (10) 1.0 (4) 0.6 (12) 90.3 (361) 0.0 (0) 0.0 (0) 7.8 (405) 35.2 Haemaphysalis flava Larvae 0.0 (0) 1.5 (12) 0.0 (0) 0.3 (6) 54.8 (219) 0.0 (0) 0.0 (0) 4.6 (237) 38.1 Nymphs 2.8 (44) 18.3 (146) 11.5 (46) 4.7 (93) 2.5 (10) 0.0 (0) 0.0 (0) 6.5 (339) 54.4 Adults 0.5 (8) 3.6 (29) 0.8 (3) 0.3 (6) 0.3 (1) 0.0 (0) 0.0 (0) 0.9 (47) 7.5 Subtotal 3.3 (52) 23.4 (187) 12.3 (49) 5.3 (105) 57.5 (230) 0.0 (0) 0.0 (0) 12.0 (623) 54.1 Ixodes nipponensis Larvae 0.0 (0) 0.0 (0) 0.0 (0) 0.9 (18) 15.0 (18) 0.0 (0) 0.0 (0) 1.5 (78) 62.9 Nymphs 0.0 (0) 2.3 (18) 3.8 (15) 0.4 (8) 0.3 (1) 0.0 (0) 0.0 (0) 0.8 (42) 33.9 Adults 0.2 (3) 0.1 (1) 0.0 (0) 0.0 (0) 0.0 (0) 0.0 (0) 0.0 (0) 0.1 (4) 3.2 Subtotal 0.2 (3) 2.4 (19) 3.8 (15) 1.3 (26) 15.3 (61) 0.0 (0) 0.0 (0) 2.4 (124) 10.7 Total mixed forests Larvae 0.0 (0) 1.5 (12) 0.0 (0) 1.3 (25) 154.8 (619) 0.0 (0) 0.0 (0) 12.6 (656) 56.9 Nymphs 3.9 (62) 21.8 (174) 16.0 (64) 5.2 (103) 5.8 (23) 0.0 (0) 0.0 (0) 8.2 (426) 37.0 Adults 0.7 (11) 3.8 (30) 1.0 (4) 0.8 (15) 2.5 (10) 0.0 (0) 0.0 (0) 1.3 (70) 6.1 Total 4.6 (73) 27.0 (216) 17.0 (68) 7.2 (143) 163.0 (652) 0.0 (0) 0.0 (0) 22.2 (1,152) 100.0 Total (n = 272) Haemaphysalis longicornis Larvae 1.2 (37) 3.0 (121) 0.7 (30) 9.4 (583) 61.9 (2,477) 268.4 (5,367) 7.2 (258) 32.6 (8,873) 59.1 Nymphs 52.4 (1,571) 17.7 (708) 49.8 (2,191) 10.8 (669) 3.0 (118) 1.3 (25) 0.0 (0) 19.4 (5,282) 35.2 Adults 0.1 (4) 1.0 (38) 5.7 (251) 7.1 (443) 3.2 (126) 0.2 (3) 0.0 (0) 3.2 (865) 5.7 Subtotal 53.7 (1,612) 21.7 (867) 56.2 (2,472) 27.3 (1,695) 68.0 (2,721) 269.8 (5,395) 7.2 (258) 55.2 (15,020) 75.8 Haemaphysalis flava Larvae 0.0 (0) 0.3 (12) 0.0 (0) 1.0 (59) 59.8 (2,393) 1.6 (31) 7.4 (268) 10.2 (2,763) 71.1 Nymphs 2.3 (70) 8.7 (348) 3.0 (134) 2.2 (135) 0.9 (34) 0.3 (6) 7.4 (266) 3.7 (993) 25.5 Adults 0.3 (10) 1.3 (52) 0.3 (13) 0.2 (14) 0.5 (18) 0.2 (3) 0.6 (23) 0.5 (133) 3.4 Subtotal 2.7 (80) 10.3 (412) 3.3 (147) 3.4 (208) 61.1 (2,445) 2.0 (40) 15.5 (557) 14.3 (3,889) 19.6 Ixodes nipponensis Larvae 0.0 (0) 0.0 (0) 0.0 (0) 7.9 (487) 6.4 (257) 0.4 (7) 0.2 (7) 2.8 (758) 83.1 Nymphs 0.2 (7) 1.1 (45) 1.0 (43) 0.2 (12) 0.3 (10) 0.1 (1) 0.1 (1) 0.4 (119) 13.1 Adults 0.2 (6) 0.3 (13) 0.3 (11) 0.1 (2) 0.0 (0) 0.0 (0) 0.1 (3) 0.1 (35) 3.8 Subtotal 0.4 (13) 1.5 (58) 1.2 (54) 8.1 (501) 6.7 (267) 0.4 (8) 0.3 (11) 3.4 (912) 4.6 Grand total Larvae 1.2 (37) 3.3 (133) 0.7 (30) 18.2 (1,129) 128.2 (5,127) 270.3 (5,405) 14.8 (533) 45.6 (12,394) 62.5 Nymphs 54.9 (1,648) 27.5 (1,101) 53.8 (2,368) 13.2 (816) 4.1 (162) 1.6 (32) 7.4 (267) 23.5 (6,394) 32.3 Adults 0.7 (20) 2.6 (103) 6.3 (275) 7.4 (456) 3.6 (144) 0.3 (6) 0.7 (26) 3.8 (1,033) 5.2 Total 56.8 (1,705) 33.4 (1,337) 60.8 (2,673) 38.8 (2,404) 135.8 (5,433) 272.2 (5,443) 22.9 (826) 72.9 (19,821) 100.0
Table 1. Monthly mean number (total) larvae, nymph, and adult ixodid ticks collected using tick drag and sweep methods for four habitats at US-ROK operated military training sites and private lands located near the DMZ, Republic of Korea, 2004 and 2005

Total number of replicates by 100 m transect for each habitat.

Table 1.

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