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Original Article

Echinostoma miyagawai Ishii, 1932 (Echinostomatidae) from Ducks in Aceh Province, Indonesia with Special Reference to Its Synonymy with Echinostoma robustum Yamaguti, 1935

Jong-Yil Chai1,2orcid, Bong-Kwang Jung1,*orcid, Taehee Chang1, Hyejoo Shin1, Jaeeun Cho1, Jin-Youp Ryu3, Hyun-Seung Kim3, Kwanghoon Park3, Mun-Hyoo Jeong3, Eui-Hyug Hoang3, Marzuki Bin Muhammad Abdullah4
The Korean Journal of Parasitology 2021;59(1):35-45.
Published online: February 19, 2021

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*Corresponding author (mulddang@snu.ac.kr)
• Received: November 11, 2020   • Revised: November 16, 2020   • Accepted: November 16, 2020

Copyright © 2021 by The Korean Society for Parasitology and Tropical Medicine

This is an Open Access article distributed under the terms of the Creative Commons Attribution Non-Commercial License (https://creativecommons.org/licenses/by-nc/4.0) which permits unrestricted non-commercial use, distribution, and reproduction in any medium, provided the original work is properly cited.

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Echinostoma miyagawai Ishii, 1932 (Echinostomatidae) from Ducks in Aceh Province, Indonesia with Special Reference to Its Synonymy with Echinostoma robustum Yamaguti, 1935
Korean J Parasitol. 2021;59(1):35-45.   Published online February 19, 2021
DOI: https://doi.org/10.3347/kjp.2021.59.1.35
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Echinostoma miyagawai Ishii, 1932 (Echinostomatidae) from Ducks in Aceh Province, Indonesia with Special Reference to Its Synonymy with Echinostoma robustum Yamaguti, 1935
Korean J Parasitol. 2021;59(1):35-45.   Published online February 19, 2021
DOI: https://doi.org/10.3347/kjp.2021.59.1.35
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Echinostoma miyagawai Ishii, 1932 (Echinostomatidae) from Ducks in Aceh Province, Indonesia with Special Reference to Its Synonymy with Echinostoma robustum Yamaguti, 1935
Image Image Image
Fig. 1 Echinostoma miyagawai from a Pitalah duck in Aceh Province, Indonesia. (A) An adult worm showing its characteristic head collar with collar spines, oral and ventral suckers, uterus with eggs, cirrus sac, ovary, Mehlis’ gland, testes, and vitelline follicles. Acetocarmine-stained. Ventral view. Scale bar=0.72 mm. (B) Head collar equipped with 37 collar spines. Acetocarmine-stained. Ventral view. Scale bar=100 μm. (C) Line drawing of the head collar with 37 collar spines. Scale bar=100 μm.
Fig. 2 A phylogenetic tree of Echinostoma miyagawai in comparison with other 37-collar-spined ‘E. revolutum group’ constructed based on 197 bp of mitochondrial cox1 sequences. ●, our specimens.
Fig. 3 A phylogenetic tree of Echinostoma miyagawai in comparison with other 37-collar-spined ‘E. revolutum group’ constructed based on 401 bp of mitochondrial nad1 sequences. ●, our specimens.

Fig. 1

Fig. 2

Fig. 3

Echinostoma miyagawai Ishii, 1932 (Echinostomatidae) from Ducks in Aceh Province, Indonesia with Special Reference to Its Synonymy with Echinostoma robustum Yamaguti, 1935

Measurements of Echinostoma miyagawai adults in comparison with other 37-collar-spined Echinostoma species (unit: μm)

Species E. miyagawai (this study) (n=14) E. miyagawai [15] (n=13) E. revolutum [15] (n=16) E. paraulum [15] (n=10)
Organ Mean Range Mean Range Mean Range Mean Range
Body length (BL) 7,186 6,186–8,536 9,990 9,163–11,014 10,531 9,454–11,846 6,345 5,600–6,862
Body width 1 (BW1) 843 868–1,274 1,159 1,029–1,272 1,298 1,182–1,454 1,043 923–1,108
Body width 2 (BW2) 1,064 667–1,053 1,026 898–1,178 1,618 1,303–1,815 1,308 1,138–1,446
Body width 3 (BW3) 1,220 1,012–1,416 1,305 1,197–1,459 1,815 1,454–2,092 1,514 1,231–1,662
Head collar length (CL) 485 395–599 434 393–505 392 351–424 488 432–553
Head collar width (CW) 614 528–725 656 598–692 607 552–652 729 644–781
Oral sucker length (OSL) 267 227–307 305 262–337 316 261–358 287 265–318
Oral sucker width (OSW) 301 233–412 293 262–355 282 246–303 335 280–371
Angle spine length (ASL) 78 71–83 70 52–93 74 58–92 100 73–124
Angle spine with (ASW) 21 19–25 22 17–29 17 11–24 21 15–28
Lateral spine length (LSL) 81 75–91 76 67–87 79 64–91 101 85–120
Lateral spine width (LSW) 21 18–24 23 20–26 17 12–21 21 17–25
Dorsal spine length (DSL) 76 69–79 68 58–75 85 71–94 99 90–105
Dorsal spine width (DSW) 21 18–24 21 19–26 18 13–23 21 19–23
Prepharynx length (PL) 18 4–41 10 0–37 41 7–99 45 15–68
Pharynx length (PHL) 273 230–292 299 262–337 237 209–258 324 273–379
Pharynx width (PHW) 277 197–335 250 187–355 219 194–258 238 212–280
Esophagus length (ESL) 659 569–790 585 542–655 408 318–569 408 265–546
Cirrus sac length (CSL) 491 410–710 506 468–598 552 410–735 413 318–531
Cirrus sac width (CSW) 262 242–292 300 281–355 316 243–403 281 205–326
Seminal vesicle length (SVL) 244 205–330 - - 336 224–440 238 152–303
Seminal vesicle width (SVW) 133 82–183 - - 146 60–212 134 114–144
Ventral sucker length (VSL) 658 594–830 706 655–748 873 796–1,038 667 569–766
Ventral sucker width (VSW) 648 544–910 738 655–785 893 796–1,061 731 705–766
Ovary length (OVL) 281 198–342 283 243–355 349 288–394 150 114–182
Ovary width (OVW) 410 290–620 397 337–449 416 291–493 246 212–288
Mehlis’ gland length (MEL) 631 413–765 386 337–468 399 326–455 153 106–190
Mehlis’ gland width (MEW) 233 135–295 373 318–411 734 582–849 310 258–341
Ant. testis length (ATL) 534 403–653 564 430–692 788 629–932 384 341–455
Ant. testis width (ATW) 728 588–982 409 337–505 597 425–750 377 326–447
Post. testis length (PTL) 564 477–674 562 449–655 879 627–1,061 406 296–515
Post. testis width (PTW) 708 593–787 411 337–524 579 395–705 415 288–531
Egg length (EL) 93 79–105 95 94–96 114 108–125 113 104–122
Egg width (EW) 62 56–70 60 59–60 65 57–75 62 53–70
Forebody length (FORE) 1,121 912–1,269 1,408 1,253–1,533 1,369 1,200–1,662 1,218 985–1,600
ODIV 982 808–1,131 1,181 1,103–1,324 1,027 923–1,262 1,074 923–1,354
OVAR 3,923 3,536–4,311 4,856 4,507–5,348 4,461 3,969–5,046 3,043 2,739–3,385
TEND 2,497 2,245–2,795 3,081 2,824–3,329 3,653 3,000–4,308 2,059 1,262–2,431
OSW/PHW 1.09 0.88–1.30 0.97 0.78–1.11 1.29 1.15–1.54 1.41 1.24–1.58
BW1/BL (%) 11.7 9.6–14.0 11.6 11.2–12.5 12.4 11.2–14.3 16.5 15.0–19.8
BW2/BL (%) 14.8 13.1–17.0 10.3 9.7–10.9 15.4 13.8–17.4 20.7 18.9–24.7
BW3/BL (%) 17.1 14.2–20.9 13.1 12.4–13.6 17.3 15.2–19.5 23.9 21.5–26.9
FORE/BL (%) 15.6 14.0–17.2 14.1 13.4–14.9 13.0 10.9–14.8 19.3 15.5–26.3
CW/BW3 (%) 50.6 43.1–58.4 50.4 46.2–56.9 33.6 28.3–38.3 48.4 42.0–57.3
OVID/BL (%) 13.7 12.3–14.7 11.8 11.2–12.5 9.8 8.3–11.2 17.0 14.5–23.7
OVAR/BL (%) 54.7 50.3–57.7 48.6 47.7–49.7 42.5 39.1–46.2 48.1 43.5–57.0
TEND/BL (%) 34.8 31.2–36.7 30.9 30.1–31.9 34.6 31.4–37.4 32.3 22.0–35.4

ODIV, distance from anterior extremity to intestinal bifurcation; OVAR, distance from the posterior margin of ventral sucker to ovary; TEND, length of post-testicular region [15].

Sequence comparison of our samples with other 37-collar-spined Echinostoma spp. based on cox1 and nad1 region

cox1 nad1
Our samples (n=4)
99.0–100		 Our samples (n=4)
99.5–100
E. miyagawai (MN116740; China; Fu et al.) 99.0–100 E. miyagawai (KP065635; Czech Republic; Georgieva et al.) 99.3–99.8
E. miyagawai (NC039532; China; Li et al.) 99.0–100 E. robustum (LC224091; Bangladesh; Mohanta et al.) 99.3–99.8
E. miyagawai (KP455602; Thailand; Nagataki et al.) 98.6–100 E. miyagawai (KP455620; Thailand; Nagataki et al.) 99.0–99.6
E. miyagawai (GU324943; Thailand; Saijuntha et al.) 98.8–100 E. friedi (AJ564379; Spain; Marcilla et al.) 99.3–99.8
E. revolutum American lineage (GQ463016; USA; Detwiler et al.) 92.6–93.2 E. revolutum (AF026287; Australia; Morgan and Blair) 98.5–99.0
E. revolutum Southeast Asian lineage (GU324945; Thailand; Saijuntha et al.) 90.4–91.6 E. miyagawai (KY436400; New Zealand; Georgieva et al.) 97.3–97.8
E. robustum (GQ463053; USA; Detwiler et al.) 94.3–94.8
E. robustum (GQ463055; Brazil; Detwiler et al.) 90.5–91.0
E. revolutum American lineage (GQ463061; USA; Detwiler et al.) 88.8
Table 1 Measurements of Echinostoma miyagawai adults in comparison with other 37-collar-spined Echinostoma species (unit: μm)

ODIV, distance from anterior extremity to intestinal bifurcation; OVAR, distance from the posterior margin of ventral sucker to ovary; TEND, length of post-testicular region [15].

Table 2 Sequence comparison of our samples with other 37-collar-spined Echinostoma spp. based on cox1 and nad1 region

Table 1

Table 2

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